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Prof. Dr. Martin Entling

Head (Ecosystem Analysis)
Visiting hours: Wed. 11-12 a.m.
Building I, room 3.08


  • Trophic interactions
  • Landscape ecology
  • Ecosystem services
  • Biodiversity
  • Spider Ecology

Curriculum vitae

2000 Diploma in Biology, University of Göttingen
2004 PhD in Agricultural Sciences, University of Göttingen
2004-2010 Research and teaching at University of Bern
2009 CAS in Higher Education, University of Bern
2010 Venia Docendi in Ecology, University of Bern
2014-2018 Head of the Institute for Environmental Sciences
since 2010 Head of Ecosystem Analysis lab at University of Koblenz-Landau




Fusser MS, Pfister SC, Entling MH & Schirmel J (2017) Effects of field margin type and landscape composition on predatory carabids and slugs in wheat fields. Agriculture, Ecosystems and Environment 247: 182-188.

Graf N, Bucher R, Schaefer RB & Entling MH (2017) Contrasting effects of aquatic subsidies on a terrestrial trophic cascade. Biology Letters 13: 20170129.

Hoffmann C, Köckerling J, Biancu S, Gramm T, Michl G & Entling MH (2017) Can flowering greencover crops promote biological control in German vineyards? Insects 8: 121.

Hudson L, Newbold T, Contu S, Hill SLL, Lysenko I, ..., Entling MH, ..., Ewers RM, Mace GM, Purves DW, Scharlemann JPW & Purvis A (2017) The database of the PREDICTS (Projecting Responses of Ecological Diversity in Changing Terrestrial Systems) project. Ecology and Evolution 7: 145-188.

Pennington T, Kraus C, Alakina E, Entling MH & Hoffmann C (2017) Minimal pruning and reduced plant protection promote predatory mites in grapevine. Insects 8: 86.

Pfister SC, Sutter L, Albrecht M, Marini S, Schirmel J & Entling MH (2017) Positive effects of local and landscape features on predatory flies in Europen agricultural landscapes. Agriculture, Ecosystems and Environment 239: 283-292.

Pfister SC, Eckerter PW, Schirmel J, Cresswell JE & Entling MH (2017) Sensitivity of commercial pumpkin yield to potential decline among different groups of pollinating bees. Royal Society Open Science 4: 170102.

Pfister SC, Schirmel J & Entling MH (2017) Aphids and their enemies in pumpkin respond differently to management, local and landscape features. Biological Control 115: 37-45.

Stiegel S, Entling MH & Mantilla-Contreras J (2017) Reading the leaves’ palm: leaf traits and herbivory along the microclimatic gradient of forest layers. PLOS ONE 12: e0169741.

Wendland S, Baudy P, Fusser MS, Entling MH & Schirmel J (2017) Contrasting predation pressure of different ground beetles on slugs and their eggs. Biocontrol 62: 749-756.



Binz H, Kraft EF, Entling MH & Menzel F (2016) Behavioral response of a generalist predator to chemotactile cues of two taxonomically distinct prey species. Chemoecology 26: 153-162.

Bucher R, Wedel MF, Entling MH, Andres C & Nickel H (2016) Biodiversity in low-intensity pastures, straw meadows, and fallows of a fen area – a multitrophic comparison. Agriculture, Ecosystems and Environment 219: 190-196.

Buri P, Humbert J-Y, Stanska M, Hajdamowicz I, Tran E, Entling MH & Arlettaz R (2016) Delayed mowing promotes planthoppers, leafhoopers and spiders in extensively managed meadows. Insect Conservation and Diversity 9: 536-545.

Buse J, Entling MH, Ranius T & Assmann T (2016) Response of saproxylic beetles to small-scale habitat connectivity depends on trophic levels. Landscape Ecology 31: 939-949.

Coudrain V, Rittiner S, Herzog F, Tinner W & Entling MH (2016) Landscape distribution of food and nesting sites affect larval diet and nest size, but not abundance of Osmia bicornis. Insect Science 23: 746-753.

De Palma A, Abrahamczyk S, Albrecht M, Baset Y, Blake RJ, Bugter R, Connop S, Cunnigham SA, Darvill B, Diekötter T, Downing N, Entling MH, ... & Purvis A (2016) ) Predicting bee community responses to land-use changes: Effects of geographic and taxonomic biases. Scientific Reports 6: 31153.

Fusser M, Pfister S, Entling MH & Schirmel J (2016) Effects of landscape composition on carabids and slugs in herbaceous and woody field margins. Agriculture, Ecosystems and Environment 226: 79-87.

Holland JM, Bianchi FJJA, Entling MH, Moonen A-C, Smith BM & Jeanneret P (2016) Structure, function and management of semi-natural habitats for conservation biological control: A review of European studies. Pest Management Science 72: 1638-1651.

Müller IB, Buhk C, Alt M, Entling MH & Schirmel J (2016) Plant functional shifts in Central European grassland under traditional flood irrigation. Applied Vegetation Science 19: 122-131.

Müller IB, Buhk C, Lange D, Entling MH & Schirmel J (2016) Contrasting effects of irrigation and fertilization on plant diversity in hay meadows. Basic and Applied Ecology 17: 576-585.

Picchi M, Bocci G, Petacchi R & Entling MH (2016) Effects of local and landscape factors on spiders and olive fruit flies. Agriculture, Ecosystems and Environment 222: 138-147.

Rader R, Bartomeus I, Garibaldi LA, Garratt MPD, Howlett B , Cunningham SA, Mayfield MM, Andersson GKS, Blanche R, Bommarco R, Brittain C, Carvalheiro LG, Chacoff NP, Entling MH, … and Woyciechowski M (2016) Non-bee insects are important contributors to global crop pollination. PNAS 113: 146-151.

Schirmel J, Bundschuh M, Entling MH, Kowarik I & Buchholz S (2016) Invasive plant effects on resident animals across ecosystems, taxa, and feeding types. A meta-analysis. Global Change Biology 22: 594-603.

Schirmel J, Thiele J, Entling MH & Buchholz S (2016) Trait composition and functional diversity of spiders and carabids in linear landscape elements. Agriculture, Ecosystems and Environment 235: 318-328.

Tschumi M, Albrecht M, Bärtschi C, Collatz J, Entling MH & Jacot K (2016) Perennial, species-rich wildflower strips enhance pest control and crop yield. Agriculture, Ecosystems and Environment 220: 97-103.

Tschumi M, Albrecht M, Collatz J, Dubsky V, Entling MH, Najar-Rodriguez AJ & Jacot K (2016) Tailored flower strips promote natural enemy biodiversity and enhance pest control in adjacent potato crops. Journal of Applied Ecology 53: 1169-1176.

Wagner J, Entling MH & Schirmel J (2016) Heuschrecken in biologisch und konventionell genutzten Weinbergen in der Pfalz. Articulata 31: 13-22.



Anjum-Zubair M, Entling MH, Bruckner A, Drapela T & Frank T (2015) Differentiation of carabid beetle assemblages between semi-natural habitats and adjoining winter wheat. Agricultural and Forest Entomology 17: 355-365.

Bucher R, Heinrich H & Entling MH (2015) Plant choice, herbivory and weight gain of wood crickets under the risk of predation. Entomologia Experimentalis et Applicata 155: 148-153.

Bucher R, Menzel F & Entling MH (2015) Impact of spider predation risk on arthropod communities and herbivory in the field. Oecologia 178: 571-577.

Buse J, Slachta M, Sladecek F, Carpaneto G & Entling MH (2015) Relative importance of pasture size and grazing continuity for the long-term conservation of European dung beetles. Biological Conservation 187: 112-119.

Buse J, Fassbender S, Entling MH & Pavlicek T (2015) Microclimatic divergence in a Mediterranean canyon affects richness, composition, and body size in saproxylic beetle assemblages. PLOS ONE 10: e0129323.

Hermann JD, Opatovsky I, Lubin Y, Pluess T, Gavish-Regev E & Entling MH (2015) Effects of non-native Eucalyptus plantations on epigeal spider communities in the northern Negev desert, Israel. Journal of Arachnology 43: 101-106.

Krell B, Röder N, Link M, Gergs R, Entling MH & Schäfer RB (2015) Aquatic prey subsidies to riparian spiders in a stream with different land use types. Limnologica 51: 1-7.

Pfister SC, Schäfer RB, Schirmel J & Entling MH (2015) Effects of hedgerows and riparian margins on aerial web-building spiders in cereal fields. Journal of Arachnology 43: 400-405.

Schulz R, Bundschuh M, Gergs R, Brühl CA, Diehl D, Entling MH, Fahse L, Frör O, Jungkunst HF, Lorke A, Schäfer RB, Schaumann GE & Schwenk K (2015) Review on environmental alterations propagating from aquatic to terrestrial ecosystems. Science of the Total Environment 538: 246-261.

Tschumi M, Albrecht M, Entling MH & Jacot K (2015) High effectiveness of tailored flower strips in reducing pests and crop plant damage. Proceedings of the Royal Society of London, Series B: Biological Sciences 282: 20151369.

Uhl P, Bucher R, Schäfer RB, Entling MH (2015) Sublethal effects of Imidacloprid in a tritrophic system. Chemosphere 132: 152-158.



Binz H, Bucher R, Entling MH & Menzel F (2014) Knowing the risk: crickets distinguish between spider predators of different size and commonness. Ethology 120: 99-110.

Bucher R, Binz H, Menzel F & Entling MH (2014) Spider cues stimulate feeding, weight gain and survival of crickets. Ecological Entomology 39: 667-673.

Bucher R, Binz H, Menzel F & Entling MH (2014) Effects of spider chemotactile cues on arthropod behaviour. Journal of Insect Behavior 27: 567-580.

Coudrain V, Schüepp C, Herzog F, Albrecht M & Entling MH (2014) Habitat amount modulates the effect of patch isolation on host-parasitoid interactions. Frontiers in Environmental Science 2: 27.

Hudson L, Newbold T, Contu S, Hill SLL, Lysenko I, ..., Entling MH, ..., Ewers RM, Mace GM, Purves DW, Scharlemann JPW & Purvis A (2014) The PREDICTS database: a global database of how local terrestrial biodiversity responds to human Impacts. Ecology and Evolution 4: 4701-4735.

Mestre L, Bucher R & Entling MH (2014) Trait-mediated effects between predators: ant chemical cues induce spider dispersal. Journal of Zoology 293: 119-125.

Schirmel J, Alt M, Rudolph I & Entling MH (2014). Effects of traditional flood irrigation on invertebrates in lowland meadows. PLOS ONE 9: e110854.

Schüepp C, Herzog F & Entling MH (2014) Disentangling multiple drivers of pollnation in a landscape-scale experiment. Proceedings of the Royal Society of London, Series B: Biological Sciences 281: 20132667.

Schüepp C, Uzman D, Herzog F & Entling MH (2014) Habitat isolation affects plant-herbivore-enemy interactions on cherry trees. Biological Control 71: 56-64.



Birkhofer K, Entling MH & Lubin Y (2013) Agroecology: Trait composition, spatial relationships, trophic interactions. Spider Research in the 21st Century (ed. D Penney), pp. 200-228. Siri Scientific Press, Manchester.

Buse J, Heinrich C, Weiter S & Entling MH (2013) Die Dungkäfer einer neu eingerichteten Waldweide bei St. Martin in der Pfalz. Mainzer Naturwissenschaftliches Archiv 50: 361-369.

Coudrain V, Herzog F & Entling MH (2013) Effects of habitat fragmentation on abundance, larval food and parasitism of a spider-hunting wasp. PLOS ONE 8: e59286.



Schüepp C, Rittiner S & Entling MH (2012) High bee and was diversity in a heterogeneous tropical farming system compared to protected forest. PLOS ONE 7: e52109.

Entling MH, Schweiger O, Bacher S, Espadaler X, Hickler T, Kumschick S, Woodcock BA & Nentwig W (2012) Species richness-environment relationships of European arthropods at two spatial grains: habitats and countries. PLOS ONE 7: e45875.

Entling MH (2012) Book Review: Invasive Alien Arthropod Predators and Parasitoids: An Ecological Approach (eds. H Roy, P De Clercq, L-J Lawson Handley, JJ Sloggett, RL Poland & E Wajnberg) Basic and Applied Ecology 13: 735-736.

Hermann JD, Kormann U, Schüepp C, Stocker Y, Herzog F & Entling MH (2012) Effects of habitat isolation and predation pressure on an arboreal food-web. Community Ecology 13: 82-87.



Bucher R & Entling MH (2011) Contrasting effects of habitat fragmentation, population density, and prey availability on body condition of two orb-weaving spiders. Ecological Entomology 36: 680-685.

Entling MH, Stämpfli K & Ovaskainen O (2011) Increased propensity for aerial dispersal in disturbed habitats due to intraspecific variation and species turnover. Oikos 120: 1099-1109.

Kumschick S, Fronzek S, Entling MH & Nentwig W (2011) Rapid spread of the wasp spider Argiope bruennichi across Europe: a consequence of climate change? Climatic Change 109: 319-329.

Sanders D & Entling MH (2011) Large variation of suction sampling efficiency depending on arthropod groups, species traits and habitat properties. Entomologia Experimentalis et Applicata 138: 234-243.

Schüepp C, Herrmann JD, Herzog F & Schmidt-Entling MH (2011) Differential effects of habitat isolation and landscape composition on wasps, bees, and their enemies. Oecologia 165: 713-721.

Stutz S & Entling MH (2011) Effects of the landscape context on aphid-ant-predator interactions on cherry trees. Biological Control 57: 37-43.


Hermann JD, Bailey D, Hofer G, Herzog F & Schmidt-Entling MH (2010) Spiders associated with the meadow and tree canopies of orchards respond differently to habitat fragmentation. Landscape Ecology 25: 1375-1384.

Bailey D, Schmidt-Entling MH, Eberhart P, Herrmann JD, Hofer G, Kormann U & Herzog F (2010) Effects of habitat amount and isolation on biodiversity in fragmented traditional orchards. Journal of Applied Ecology 47: 1003-1013.

Mühlner S, Kormann U, Schmidt-Entling MH, Herzog F & Bailey D (2010) Structural versus functional habitat connectivity measures to explain bird diversity in fragmented orchards. Journal of Landscape Ecology 3: 52-63.

Anjum-Zubair M, Schmidt-Entling MH, Querner P & Frank T (2010) Influence of within-field position and adjoining habitat type on carabid beetle assemblages in winter wheat. Agricultural and Forest Entomology 12: 301-306.

Bruggisser O, Schmidt-Entling MH & Bacher S (2010) Effects of vineyard management on biodiversity at three trophic levels. Biological Conservation 143: 1521-1528.

Entling W, Schmidt-Entling MH, Bacher S, Brandl R & Nentwig W (2010) Body size - climate relationships of European spiders. Journal of Biogeography 37: 233-237.

Pluess T, Opatovski I, Gavish-Regev E, Lubin Y & Schmidt-Entling MH (2010) Non-crop habitats in the landscape enhance spider diversity in wheat fields of a desert agroecosystem. Agriculture, Ecosystems and Environment 137: 68-74.

Opatovski I, Plüss T, Schmidt-Entling MH, Gavish-Regev E & Lubin Y (2010) Are spider assemblages in fragmented, semi-desert habitat affected by increasing cover of agricultural crops? Agriculture, Ecosystems and Environment 135: 233-237.



Kumschick S, Schmidt-Entling MH, Bacher S, Hickler T, Entling W & Nentwig W (2009) Water limitation prevails over energy in European diversity gradients of sheetweb spiders (Araneae: Linyphiidae). Basic and Applied Ecology 10: 754-762.

Kumschick S, Schmidt-Entling MH, Bacher S, Hickler T, Espadaler X & Nentwig W (2009) Determinants of local ant (Hymenoptera: Formicidae) species richness and activity density across Europe. Ecological Entomology 34: 748-754.

Schmidt-Entling MH & Siegenthaler E (2009) Herbivore release through cascading risk effects. Biology Letters 5: 773-776.

Farwig N, Bailey D, Bochud E, Herrmann JD, Kindler E, Reusser N, Schuepp C & Schmidt-Entling MH (2009) Isolation from forest reduces pollination, seed predation and insect scavenging in Swiss farmland. Landscape Ecology 24: 919-927.

Schmidt-Entling MH & Döbeli J (2009) Sown wildflower areas to enhance spiders in arable fields. Agriculture, Ecosystems and Environment 133: 19-22.

Eichenberger B, Siegenthaler E & Schmidt-Entling MH (2009) Body size determines the outcome of competition for webs among alien and native sheetweb spiders (Araneae: Linyphiidae). Ecological Entomology 34: 363-368.

Muff P, Kropf C, Frick H, Nentwig W & Schmidt-Entling MH (2009) Coexistence of divergent communities at natural boundaries: spider (Arachnida: Araneae) diversity across an alpine timberline. Insect Conservation and Diversity 2: 36-44.


Jud P & Schmidt-Entling MH (2008) Fluid type, dilution, and bitter agent influence spider preservation in pitfall traps. Entomologia Experimentalis et Applicata 129: 356-359.

Schmidt MH, Rocker S, Hanafi J & Gigon A (2008) Rotational fallows as overwintering habitat for grassland arthropods: the case of spiders in fen meadows. Biodiversity and Conservation 17: 3003-3012.

Pluess T, Opatovsky I, Gavish-Regev E, Lubin Y & Schmidt MH (2008) Spiders in wheat fields and semi-desert in the Negev (Israel). Journal of Arachnology 36: 368-373.

Schmidt MH, Thies C, Nentwig W & Tscharntke T (2008) Contrasting responses of arable spiders to the landscape matrix at different spatial scales. Journal of Biogeography 35: 157-166.


Entling W, Schmidt MH, Bacher S, Brandl R & Nentwig W (2007) Niche properties of Central European spiders: shading, moisture, and the evolution of the habitat niche. Global Ecology and Biogeography 18: 440-448.

Muff P, Schmidt MH, Frick H, Nentwig W & Kropf C (2007) Spider (Arachnida: Araneae) distribution across the timberline in the Swiss Central Alps (Alp Flix, Grisons) and three morphologically remarkable species. Arachnologische Mitteilungen 34: 16-24.

Schmidt MH & Hänggi A (2007) Zelotes mundus (Araneae: Gnaphosidae) in the Camargue: a continental species meets the Mediterranean coast. Bulletin of the British Arachnological Society 14: 27-29.


Schmidt MH, Clough Y, Schulz W, Westphalen A & Tscharntke T (2006) Capture efficiency and preservation attributes of different fluids in pitfall traps. Journal of Arachnology 34: 159-162.

Flückiger R & Schmidt MH (2006): Contribution of sown wildflower areas to cereal aphid control: from local to landscape scale. IOBC/WPRS Bulletin 29: 41-44.


Schmidt MH, Roschewitz I, Thies C & Tscharntke T (2005) Differential effects of landscape and management on diversity and density of ground-dwelling farmland spiders. Journal of Applied Ecology 42: 281-287.

Schmidt MH & Tscharntke T (2005) Landscape context of sheetweb spider (Araneae: Linyphiidae) abundance in cereal fields. Journal of Biogeography 32: 467-473.

Schmidt MH & Tscharntke T (2005) The role of perennial habitats for Central European farmland spiders. Agriculture, Ecosystems and Environment 105: 235-242.

Schmidt MH, Lefebvre G, Poulin B & Tscharntke T (2005) Reed cutting affects arthropod communities, potentially reducing food for passerine birds. Biological Conservation 121: 157-166.


Schmidt MH, Thewes U, Thies C & Tscharntke T (2004) Aphid suppression by natural enemies in mulched cereals. Entomologia Experimentalis et Applicata 113: 87-93.

Schmidt MH, Thies C & Tscharntke T (2004) The landscape context of arthropod biological control. Ecological Engineering for Pest Management: Advances in Habitat Manipulation for Arthropods (eds. GM Gurr, SD Wratten & MA Altieri), pp. 55-63. CSIRO, Collingwood VIC.


Schmidt MH, Lauer A, Purtauf T, Thies C, Schaefer M & Tscharntke T (2003) Relative importance of predators and parasitoids for cereal aphid control. Proceedings of the Royal Society of London, Series B: Biological Sciences 270: 1905-1909.